Escharella Gray, 1848

Type species: Lepralia immersa Fleming, 1828

Synonyms Author Type Species
Haywardipora Soule, Soule & Chaney, 1995 Haywardipora rugosa Soule, Soule & Chaney, 1995
Mucronella Hincks, 1880 Lepralia peachi Johnston, 1847 (=Lepralia immersa)
Perigastrella Hincks, 1880 Discopora coccinea labiata Boeck in Smitt, 1868

This genus includes the following species. (This list is not definitive; many species originally assigned to Mucronella and Perigastrella are included here, but their identity has not been checked. Critical reviews of data are welcome.)

  1. Escharella abyssicola
    		2. (Norman, 1869) Recent NE Atlantic WoRMS 111480
  2. Escharella acuta
    		3. Zabala, Maluquer & Harmelin, 1993 Recent Mediterranean WoRMS 111481
  3. Escharella anatirostris
    		4. (O'Donoghue, 1924) Recent S. Africa WoRMS 470311
  4. Escharella anatirostris spinifera
    		5. (O'Donoghue, 1924) Recent S. Africa WoRMS 472711
  5. Escharella areolata
    		6. ?     WoRMS 1414679
  6. Escharella areolata parva
    		7. (Cipolla, 1928) Tertiary Libya WoRMS 1415150
  7. Escharella ashapuraae
    		8.  Guha & Gopikrishna, 2007 Neogene Miocene Aquitanian India Gujarat WoRMS 470320
  8. Escharella capitata
    		9. (Canu & Lecointre, 1930) Tertiary W. France WoRMS 1411008
  9. Escharella connectens
    		10. (Ridley, 1881) Recent S. Atlantic WoRMS 111482
  10. Escharella? cothurnica
    		11.  (Kirkpatrick, 1888) Recent Indian Ocean - Mauritius WoRMS 471049
  11. Escharella crozetensis
    		12. (Waters, 1904) Recent S. Indian O. WoRMS 470308
  12. Escharella cryptooecium
    		13.  Souto, Fernandez-Pulpeiro & Reverter-Gil, 2007 Recent East Atlantic WoRMS 470318
  13. Escharella cycloris
    		14.  (Gabb & Horn, 1862) Paleogene Oligocene USA Alabama WoRMS 1411009
  14. Escharella diaphana
    		15. (MacGillivray, 1879) Recent Australia WoRMS 470305
  15. Escharella dijmphnae
    		16. (Kluge, 1929) Recent Arctic WoRMS 470310
  16. Escharella discors
    		17. Hayward & Cook, 1983 Recent S. Africa WoRMS 470306
  17. Escharella? elongata
    		18.  (Canu & Bassler, 1935) Paleogene Oligocene Australia Victoria WoRMS 1411010
  18. Escharella fistula
    		19. Brown, 1952 Tertiary New Zealand WoRMS 1402046
  19. Escharella fusca
    		20. (O'Donoghue & O'Donoghue, 1926) Recent Canada WoRMS 470316
  20. Escharella gilsoni
    		21.  De Blauwe, 2006 Recent Belgium WoRMS 247907
  21. Escharella granulosa
    		22. (Canu & Bassler, 1920) Paleogene Eocene Priabonian USA Mississippi WoRMS 1411011
  22. Escharella grossa
    		23. Moissette, 1988 U. Miocene Algeria WoRMS 1402047
  23. Escharella grotriani
    		24.  (Stoliczka, 1862) Paleogene Oligocene Germany WoRMS 1411012
  24. Escharella guernei
    		25.  (Jullien & Calvet, 1903) Recent East Atlantic WoRMS 865053
  25. Escharella? hoernesi
    		26.  (Reuss, 1864) Paleogene Oligocene Germany WoRMS 1411013
  26. Escharella? hoernesi paroiporosa
    		27.  Canu & Bassler, 1929 Paleogene Eocene Lutetian Belgium WoRMS 1414802
  27. Escharella hozawai
    		28. (Okada, 1929) Recent Japan WoRMS 470314
  28. Escharella immersa
    		29. (Fleming, 1828) Recent North Atlantic, Arctic WoRMS 111484
  29. Mucronella inconspicua
    		30.  Ortmann, 1890 Recent Northwest Pacific - Japan WoRMS 471051
  30. Escharella incudifera
    		31. Gordon, 1984 Recent Kermadec Ridge WoRMS 470302
  31. Escharella indivisa
    		32. Levinsen, 1916 Recent Arctic WoRMS 111485
  32. Escharella klugei
    		33. Hayward, 1979 Recent N. Atlantic, Arctic WoRMS 111486
  33. Escharella labiata
    		34. (Boeck in Smitt, 1868) Recent Atlantic WoRMS 111487
  34. Escharella labiosa
    		35. (Busk, 1856) Recent Britain (S) WoRMS 111488
  35. Escharella lagaaiji
    		36. Buge, 1957 Tertiary W. France WoRMS 1402048
  36. Escharella laqueata
    		37. (Norman, 1864) Recent NE Atlantic WoRMS 111489
  37. Escharella latodonta
    		38. Kluge, 1962 Recent Arctic WoRMS 423358
  38. Escharella levinseni
    		39. Hayward, 1994 Recent Faroe Is. WoRMS 111490
  39. Escharella longicella
    		40.  (Canu, 1914) Paleogene Eocene Lutetian France WoRMS 1437465
  40. Escharella longicollis
    		41. (Jullien, 1882) Recent NE Atlantic WoRMS 111491
  41. Escharella lopezfei
    		42.  Souto, Berning & Ostrovsky, 2016 Recent Northeast Atlantic Spain WoRMS 868424
  42. Escharella macrodonta
    		43. Levinsen, 1917 Recent Arctic WoRMS 990200
  43. Escharella mamillata
    		44. Hayward & Thorpe, 1989 Recent Clarence Is. WoRMS 173661
  44. "Escharella” massiliana
    		45.  Harmelin & Rosso, 2023 Recent Mediterranean France WoRMS 1666963
  45. Escharella modica
    		46. (Canu, 1926) Paleogene Eocene Bartonian Belgium WoRMS 1411014
  46. Escharella nilotica
    		47. (Ziko, 1985) Eocene Lutetian-Bartonian Egypt WoRMS 1411015
  47. Escharella obscura
    		48.  Norman, 1909 Recent East Atlantic Madeira WoRMS 470317
  48. Escharella octodentata
    		49. (Hincks, 1880) Recent NE Atlantic, Mediterranean WoRMS 111492
  49. Escharella ovoidea
    		50. (Reuss, 1848) Neogene Miocene Tortonian Austria WoRMS 1411016
  50. Escharella patens
    		51. (Canu & Bassler, 1920) Paleogene Eocene Priabonian USA North Carolina WoRMS 1411017
  51. Escharella praealta
    		52. (Calvet, 1906) Recent W. Europe WoRMS 111493
  52. Escharella quadrata
    		53. lopez de la Cuadra & Garcia-Gomez, 2001 Recent Mediterranean WoRMS 221556
  53. Escharella reussiana
    		54. (Busk, 1859) Tertiary W. Europe WoRMS 1411018
  54. Escharella rogeri
    		55. David, Mongereau & Pouyet, 1972 Neogene Miocene Burdigalian S. France WoRMS 1402049
  55. Escharella rugosa
    		56. (Soule, Soule, & Chaney, 1995) Recent California WoRMS 470307
  56. Escharella rylandi
    		57. Geraci, 1974 Recent Mediterranean WoRMS 111494
  57. Escharella selseyensis
    		58. Cheetham, 1966 Eocene England WoRMS 1402050
  58. Escharella serratilabris
    		59. (O'Donoghue, 1924) Recent S. Africa WoRMS 470312
  59. Escharella similis
    		60.  Ramalho, Rodríguez-Aporta & Gofas, 2022 Recent Mediterranean Spain WoRMS 1561327
  60. Escharella spinosissima
    		61. (Hincks, 1881) Recent SE Australia, New Zealand WoRMS 470304
  61. Escharella takatukii
    		62. (Okada, 1929) Recent Japan WoRMS 470315
  62. Escharella tenera
    		63. (Reuss, 1874) Eocene Austria, Hungary WoRMS 1411019
  63. Escharella teres
    		64. (Hincks, 1881) Recent Australia WoRMS 470303
  64. Escharella thompsoni
    		65.  Kluge, 1914 Recent Antarctic WoRMS 465716
  65. Escharella trispinosa
    		66. Li, 1990 L. Miocene France WoRMS 1402051
  66. Escharella? uncifera
    		67. (Canu & Bassler, 1929) Recent Philippines WoRMS 470313
  67. Escharella variolosa
    		68. (Johnston, 1838) Recent NW Europe WoRMS 111495
  68. Escharella ventricosa
    		69. (Hassall, 1842) Recent NE Atlantic, W Mediterranean WoRMS 111496
  69. Escharella ventricosa parva
    		70. Buge, 1957 Tertiary W.France WoRMS 1414803
  70. Escharella vigneauxi
    		71. David, 1965 Tertiary France WoRMS 1402052
  71. Escharella watersi
    		72. Hayward & Thorpe, 1989 Recent Antarctica WoRMS 173663

The following species have been assigned to Mucronella, and are included here with the hope that they will be revised eventually.

  1. Mucronella beneckei
    		2.  Koschinsky, 1885 Paleogene Eocene Lutetian Germany WoRMS 1405607
  2. Mucronella formidabilis
    		3.  Thornely, 1907 Recent Indian Ocean WoRMS 990416
  3. Mucronella cystioides
    		4.  Koschinsky, 1885 Paleogene Eocene Lutetian Germany WoRMS 1405608
  4. Mucronella hians
    		5. (Hennig, 1892) Cretaceous Sweden WoRMS 1412281
  5. Mucronella obesa
    		6.  Faura & Canu, 1916 Paleogene Eocene Priabonian Spain WoRMS 1405610

The following species have been assigned to Perigastrella, and are included here with the hope that they will be revised eventually.

  1. Perigastrella ansata
    		2. Canu & Bassler, 1920 Tertiary USA WoRMS 1406355
  2. Perigastrella? costifera
    		3. Canu & Bassler, 1920 Tertiary USA WoRMS 1406356
  3. Perigastrella crassilabiata (Homonym?)
    		4.  Canu & Lecointre, 1930
    Not Duvergier, 1921    		 Tertiary W. France WoRMS 1406357
  4. Perigastrella depressa
    		5. Canu & Bassler, 1920 Tertiary USA WoRMS 1406358
  5. Perigastrella elegans
    		6. Canu & Bassler, 1920 Tertiary USA WoRMS 1406359
  6. Perigastrella exserta
    		7. (Gabb & Horn, 1862) Tertiary USA WoRMS 1412844
  7. Perigastrella granulata
    		8. Zagorsek, 1994 Eocene Austria, Hungary WoRMS 1406360
  8. Perigastrella helvetica
    		9. Canu & Lecointre, 1930 Tertiary W. France WoRMS 1406361
  9. Perigastrella hexagonalis
    		10. Canu & Bassler, 1920 Tertiary USA WoRMS 1406362
  10. Perigastrella labiatula
    		11. Canu & Lecointre, 1930 Tertiary France WoRMS 1406363
  11. Perigastrella lunata
    		12. Canu & Bassler, 1924 Miocene Hungary WoRMS 1406364
  12. Perigastrella lyrulata
    		13. Canu & Lecointre, 1930 Tertiary France WoRMS 1406365
  13. Perigastrella maxilla
    		14. Canu & Bassler, 1920 Tertiary USA WoRMS 1406366
  14. Perigastrella oscitans
    		15. Canu & Bassler, 1920 Tertiary USA WoRMS 1406367
  15. Perigastrella ovalis
    		16. Canu & Bassler, 1929 Recent Philippines WoRMS 1415320
  16. Perigastrella ovoidea
    		17. Canu & Bassler, 1917 Tertiary USA WoRMS 1406368
  17. Perigastrella? parvicella
    		18. Canu & Bassler, 1924 Miocene Hungary WoRMS 1406369
  18. Perigastrella plana
    		19. Canu & Bassler, 1920 Tertiary USA WoRMS 1406370
  19. Perigastrella rectilineata
    		20. Canu & Bassler, 1920 Tertiary USA WoRMS 1406371
  20. Perigastrella rhomboidalis
    		21. Canu & Bassler, 1920 Tertiary USA WoRMS 1406372
  21. Perigastrella serrulata
    		22. (Reuss, 1848) Tertiary Europe WoRMS 1412845
  22. Perigastrella sinedorso
    		23. Duvergier, 1921 Miocene France WoRMS 1406373
  23. Perigastrella trapezoidea
    		24. Canu & Bassler, 1920 Tertiary USA WoRMS 1406374
  24. Perigastrella? tubulosa
    		25. Canu & Bassler, 1920 Tertiary USA WoRMS 1406375
Escharella diaphana Escharella ventricosa
Escharella diaphana Escharella ventricosa

Escharella page at NMITA

Escharella immersa at Atlas of Arctic Bryozoa - Piotr Kuklinski.

Escharella macrodonta at Atlas of Arctic Bryozoa - Piotr Kuklinski.

Harmer notebook - entry for Escharella
Harmer notebook - entry for Mucronella
Harmer notebook - entry for Perigastrella

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Edited by Phil Bock
Modified on 25/01/2016
Modified on 20/4/2017
Modified on 17/9/2017
Modified on 23/6/2018 (Moved M.cretacea to Balantiostoma)
Modified on 28/5/2020 (Added WoRMS links)
Modified on 2/2/2022 (Added E.similis)
Modified on 20/6/2023 (Added E. massiliana)
This URL is http://bryozoa.net/cheilostomata/romancheinidae/escharella.html