Systematic List of Families of Bryozoa

('Corrected' terminations of some systematic names not yet included).
Many of the families and genera of cheilostomes now have pages without illustrations of species. The links leading to illustrations are marked with the symbol "†".

Index: (Jump to the appropriate taxonomic group)

Order Cyclostomata
Order(?) Hederellida
Order Cryptostomida
Order Cystoporida
Order Trepostomatida
Order Fenestrida
Order Ctenostomata
Order Cheilostomata
Suborder Flustrina
Suborder Ascophorina
Infraorder Umbonulomorpha
Infraorder Lepraliomorpha
Class Phylactolaemata

CLASS STENOLAEMATA

An alternative classification of the Stenolaemata, based upon Recent Russian systematics, is given in an alternative document. The arrangement of families in the file system, and in this page that you are reading, is not intended to imply endorsement of the systematic scheme.

ORDER CYCLOSTOMATA

Distinguishing Features
Family-level systematics of the cyclostomes is poorly known. Some families are distinctive and well defined. However, others are inadequately defined, and there is no consistent version of a classification. The following list includes most of the families included in a variety of references. Several of these are either not very useful, or are considered by most authorities to be invalid. These are indicated by a smaller font in this list.

SUBORDER PALEOTUBULIPORINA

SUBORDER TUBULIPORINA

SUBORDER FASCICULINA

SUBORDER ARTICULINA

SUBORDER CANCELLATA

SUBORDER CERIOPORINA

SUBORDER RECTANGULATA

Cyclostomatids with family placement uncertain

ORDER (?) HEDERELLIDA

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ORDER CRYPTOSTOMIDA

Distinguishing Features

Families from Boardman, R.S. et al. 1983, Treatise on Invertebrate Paleontology, Part G (Revised) Vol.1; and Taylor (1993). Further suborders and families have been added from the work of Goryunova (1985). I have included several families proposed in the past 15 years, but which appear not to have been generally adopted. I would like to hear from anyone with critical comments that can help towards a reasonably stable classification.

SUBORDER PTILODICTYINA

SUBORDER RHABDOMESINA

SUBORDER TIMANODICTYINA

SUBORDER GOLDFUSSITRYPINA

SUBORDER STREBLOTRYPINA

Unplaced Cryptostomids

ORDER CYSTOPORIDA

Distinguishing Features

Families from Boardman, R.S. et al. 1983, Treatise on Invertebrate Paleontology, Part G (Revised) Vol.1.

SUBORDER CERAMOPORINA

SUBORDER FISTULIPORINA

Unplaced Cystoporid genera

ORDER TREPOSTOMATIDA

Distinguishing Features

(Families from Taylor, 1993)

Unplaced Trepostome genera

ORDER FENESTRIDA

Distinguishing Features

(Families from Taylor, 1993)

Unplaced Fenestrate genera

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CLASS GYMNOLAEMATA

ORDER CTENOSTOMATA

Distinguishing Features

Cladistic analysis of ctenostomes strongly supports the view that this is a paraphyletic grouping, with the clade including all non-phylactolaemate bryozoans (Todd, 1990). The name Gymnolaemata is suggested (by Todd) for this clade. The term 'Ctenostomata' would thus be used for an informal, paraphyletic grouping of non-skeletized taxa. A page outlining the proposed phylogenetic classification will be prepared, as soon as I work out how to format it! For the present, the arrangement of files and directories on this site will continue using traditional hierarchical systematics.

PROTOCTENOSTOMATA

SUPERFAMILY BENEDENIPOROIDEA

SUBORDER ALCYONIDIINA

SUPERFAMILY ALCYONIDIOIDEA

SUBORDER FLUSTRELLIDRINA

SUPERFAMILY HAYWARDOZOONOIDEA

SUPERFAMILY FLUSTRELLIDROIDEA

SUBORDER VICTORELLINA

SUPERFAMILY VICTORELLOIDEA

SUBORDER PALUDICELLINA

SUPERFAMILY PALUDICELLOIDEA

SUBORDER VESICULARINA

SUPERFAMILY VESICULARIOIDEA

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SUBORDER STOLONIFERINA

SUPERFAMILY TRITICELLOIDEA

SUPERFAMILY AEVERRILLIOIDEA

SUPERFAMILY VALKERIOIDEA

SUPERFAMILY ARACHNIDIOIDEA

SUPERFAMILY TEREBRIPOROIDEA

SUPERFAMILY HISLOPIOIDEA

SUPERFAMILY PENETRANTIINA

Unplaced Ctenostomates

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ORDER CHEILOSTOMATA

Distinguishing Features

The use of the term 'Cheilostomatida' for this Order was proposed a few years ago, to achieve consistency in terminations of ordinal names. As this suggestion has not become generally adopted, the usage is not continued in these web pages. The arrangement of cheilostome families is based upon an Interim Classification by D.P. Gordon. Much of the background of this classification has not yet been published. The Working Lists are accessible as Adobe Portable Document Format (PDF) files. These are: the Systematic List of Families, and the Alphabetic List of Genera (Update 6th August 2014)
Many of the families and genera now have pages without illustrations of species. The links leading to illustrations are marked with the symbol "†".

"Anasca": The Suborder Anasca has been used widely previously, and is now considered to be an informal grouping of the Suborders Inovicellina, Scupariina, Malacostegina, and the Superfamilies Calloporoidea, Buguloidea, Flustroidea and Cellarioidea in the Flustrina. It is clear that much further work is needed on the relationships between the families of cheilostomes.

SUPERFAMILY LABIOSTOMELLOIDEA

SUBORDER INOVICELLINA

SUPERFAMILY AETEOIDEA

SUBORDER SCRUPARIINA

SUPERFAMILY SCRUPARIOIDEA

SUBORDER MALACOSTEGINA

SUPERFAMILY MEMBRANIPOROIDEA

SUBORDER TENDRINA

SUPERFAMILY TENDROIDEA

SUBORDER THALAMOPORELLINA

SUPERFAMILY THALAMOPORELLOIDEA

SUBORDER BELLULOPORINA

SUPERFAMILY BELLULOPOROIDEA

SUBORDER FLUSTRINA

(SUBORDER NEOCHEILOSTOMINA is coextensive with this current definition of Flustrina.)

SUPERFAMILY CALLOPOROIDEA

SUPERFAMILY FLUSTROIDEA

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SUPERFAMILY BUGULOIDEA

SUPERFAMILY MICROPOROIDEA

SUPERFAMILY CELLARIOIDEA

SUPERFAMILY MONOPORELLOIDEA

Unplaced Families

Unplaced Anascan genera

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"SUBORDER ASCOPHORA"

The Suborder Ascophora is now considered as an informal morphological grouping of the following superfamilies in this list.

INFRAORDER ACANTHOSTEGA

SUPERFAMILY CRIBRILINOIDEA

SUPERFAMILY BIFAXARIOIDEA

SUPERFAMILY SCORIOPOROIDEA

SUPERFAMILY CATENICELLOIDEA

INFRAORDER HIPPOTHOOMORPHA

SUPERFAMILY HIPPOTHOOIDEA

INFRAORDER UMBONULOMORPHA

SUPERFAMILY ARACHNOPUSIOIDEA

SUPERFAMILY ADEONOIDEA

SUPERFAMILY PSEUDOLEPRALIOIDEA

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SUPERFAMILY LEPRALIELLOIDEA (UMBONULOIDEA)

SUPERFAMILY CHLIDONIOPSOIDEA

INFRAORDER LEPRALIOMORPHA

SUPERFAMILY SMITTINOIDEA

SUPERFAMILY SCHIZOPORELLOIDEA

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SUPERFAMILY URCEOLIPOROIDEA

SUPERFAMILY DIDYMOSELLOIDEA

SUPERFAMILY EUTHYRISELLOIDEA

SUPERFAMILY SIPHONICYTAROIDEA

SUPERFAMILY MAMILLOPOROIDEA

SUPERFAMILY CELLEPOROIDEA

SUPERFAMILY CONESCHARELLINOIDEA

Unplaced Ascophoran genera

CLASS PHYLACTOLAEMATA

Distinguishing Features

ORDER PLUMATELLIDA

Vinogradov (2004) has proposed that the Phylactolaemata be considered as a separate phylum, within a Superphylum Phoronidomorpha. This scheme is not followed here, as the results from molecular studies do not support this. In addition, the new Family Hyalinellidae Vinogradov, 2004 is not applied here.

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Home Page Alphabetic Family List
Edited by Phil Bock
Modified on 13/12/2014
This URL is http://bryozoa.net/famsys.html